Document Type

Article

Language

eng

Format of Original

10 p.

Publication Date

8-2013

Publisher

Nature Publishing Group

Source Publication

Nature Communications

Source ISSN

2041-1723

Original Item ID

doi: 10.1038/ncomms3281

Abstract

Srs2 dismantles presynaptic Rad51 filaments and prevents its re-formation as an anti-recombinase. However, the molecular mechanism by which Srs2 accomplishes these tasks remains unclear. Here we report a single-molecule fluorescence study of the dynamics of Rad51 filament formation and its disruption by Srs2. Rad51 forms filaments on single-stranded DNA by sequential binding of primarily monomers and dimers in a 5′–3′ direction. One Rad51 molecule binds to three nucleotides, and six monomers are required to achieve a stable nucleation cluster. Srs2 exhibits ATP-dependent repetitive motion on single-stranded DNA and this activity prevents re-formation of the Rad51 filament. The same activity of Srs2 cannot prevent RecA filament formation, indicating its specificity for Rad51. Srs2’s DNA-unwinding activity is greatly suppressed when Rad51 filaments form on duplex DNA. Taken together, our results reveal an exquisite and highly specific mechanism by which Srs2 regulates the Rad51 filament formation.

Comments

Published version. Nature Communications, Vol. 4 (August 2013). DOI. © 2013 Nature Publishing Group. Used with permission.

Edwin Antony was affiliated with Utah State University at the time of publication.

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